Yongqing Bao
Overall winner. Yongqing Bao took home the top award for this picture of a standoff between a Tibetan fox and a marmot, captured at China's Qilian Mountains National Nature Reserve.
Stefan Christmann
Portfolio award. Part of his award-winning photo series, this picture by German photographer Stefan Christmann shows 5,000 male emperor penguins huddling against the wind on the sea ice of Antarctica's Atka Bay.
Shangzhen Fan
Animals in their environment. Chinese photographer Shangzhen Fan won this category for his photo of a herd of Tibetan antelope leaving a trail of footprints in the snow, on the slopes of China's Kumukuli Desert.
Ripan Biswas
Animal portraits. Ripan Biswas was photographing an ant colony in West Bengal, India, when he noticed this imposter -- a crab spider, which had been mimicking his neighbors in order to infiltrate the colony and eventually feast on its inhabitants.
Thomas Easterbrook
10 years old and younger. Young photographer Thomas Easterbrook took this remarkable picture of a hummingbird hawkmoth while on holiday with his family in France.
Cruz Erdmann
11-14 years old. Cruz Erdmann was diving off Indonesia when he came across a bigfin reef squid glowing underwater.
Riccardo Marchegiani
15-17 years old. Italian teen Riccardo Marchgiani found this female gelada monkey,, with a week-old infant clinging to her belly, climbing over a cliff edge in Ethiopia's Simien Mountains National Park.
Zorica Kovacevic
Plants and fungi. The arms of a Monterey cypress tree twist and weave outwards in California's Point Lobos State Natural Reserve, in a picture taken by Zorica Kovacevic.
Max Waugh
Black and white. Max Waugh took this picture of a lone American bison in Yellowstone National Park, during a brutal snowfall.
Audun Rikardsen
Behavior: Birds. Audun Rikardsen bolted a camera and tripod, complete with a motion sensor, to a tree branch near his home in Norway. It took three years for this golden eagle to get used to the camera and start using the branch to survey the landscape below.
Manuel Plaickner
Behaviour: Amphibians and Reptiles. Manuel Plaickner's photo shows a mass migration of frogs at a pond in South Tyrol, Italy.
Jérémie Villet
Rising Star Portfolio Award. Jérémie Villet spent a month in the middle of winter observing North American mountain sheep in Yukon, northwest Canada.
Ingo Arndt
Behavior: Mammals (joint winner). A puma attacks a guanaco in Patagonia, Chile. Ingo Arndt shared the category prize with the overall winner, Yongqing Bao, after spending seven months tracking wild pumas on foot.
David Doubilet
Underwater. David Doubilet caught this colony of garden eels, which stretched nearly to the size of a football field, deep within the famous "Coral Triangle" off the coast of the Philippines.
Charlie Hamilton James
Urban wildife. A spine-tingling city scene on Pearl Street, Lower Manhattan, shows brown rats scampering near their home under a tree grille. Charlie Hamilton James captured the group as they went looking for food on New York's sidewalks.
Alejandro Prieto
Wildlife Photojournalism: Single Image. A photo of a male jaguar is projected onto a part of the US-Mexico border fence, on a star-filled night in Arizona. The moment was captured by Alejandro Prieto.
Luis Vilariño Lopez
Earth's Environments. Lava flows into the Pacific Ocean following an eruption of Kîlauea, an active volanco on Hawaii's Big Island. This aerial image was taken by Luis Vilariño Lopez.
Jasper Doest
Wildlife Photojournalist Story Award. Riku, a Japanese macaque legally captured in the wild, has performed comedy skits three times a day to audiences at the Nikkō Saru Gundan theatre, north of Tokyo. Jasper Doest, who gained permission to photograph one of Riku's shows, was appalled that such creatures were exploited for comedy.
Daniel Kronauer
Behaviour: Invertebrates. Daniel Kronauer caught a swarm of army ants moving through a rainforest in northeastern Costa Rica.
CNN  — 

It could almost be a scene from a slapstick comedy: a marmot stands frozen in fear, slack-jawed and balanced on one foot, as it suddenly notices a charging fox.

The dramatic image, captured with perfect timing by Chinese photographer Yongqing Bao, has won the prestigious Wildlife Photographer of the Year award, given out annually by London’s Natural History Museum.

He was chosen from a longlist of various category winners, all of whom managed to record the oddities, rivalries and beauty of the natural world.

Bao caught the scene on the snow-draped sloped of China’s Qilian Mountains in early spring.

He had been observing the interactions between the two creatures for some time; around an hour earlier, the marmot had spotted the fox and raised the alarm to his neighbors to get underground.

But the fox lay low and still; and, believing the coast was clear, the marmot eventually emerged to find food.

In an instant the fox dashed forward and, thanks to some lightning-quick reflexes, Bao was able to immortalize a frightening moment of realization as the marmot comes face to face with its mortality.

“Photographically, it is quite simply the perfect moment,” said Roz Kidman Cox, the chair of the judging panel, in a statement. “The expressive intensity of the postures holds you transfixed, and the thread of energy between the raised paws seems to hold the protagonists in perfect balance.”

“Images from the Qinghai-Tibet Plateau are rare enough, but to have captured such a powerful interaction between a Tibetan fox and a marmot – two species key to the ecology of this high-grassland region – is extraordinary,” Cox added.

“This compelling picture captures nature’s ultimate challenge – its battle for survival,” Natural History Museum director Michael Dixon added.

Bao collected the award at a ceremony, held at the London museum on Tuesday evening.