CNN  — 

A team of leading art researchers has revealed that two bronze sculptures of nude men riding panther-like creatures are the work of Michelangelo, bearing hallmarks of the Renaissance artist’s oeuvre including distinctive toes and his signature “eight-pack” stomach muscles.

The team, led by the Fitzwilliam Museum in Cambridge, England, conducted close to four years of research to confirm a claim from February 2015 that the sculptures were Michelangelo’s only surviving bronzes.

The museum released extensive evidence Wednesday confirming the attribution of the sculptures, known as the Rothschild bronzes, highlighting their unique anatomical features.

Researchers employed the expertise of scientists – including Peter Abrahams, professor of clinical anatomy at Warwick Medical School – to decipher key characteristics of the master’s works.

JACK TAYLOR/AFP/Getty Images
Researchers highlighted distinctive anatomical features in the sculptures, including crooked toes and 'eight-pack' torsos.

Abrahams noted that while the statues took his breath away, he was focused on unveiling “anatomical, scientific accuracy.”

“Being an observant person, both as a doctor and a scientist, I noticed that the toes on the bronzes were a bit off,” the Daily Telegraph reported him as saying.

“I then went and had a look at all the toes that I could find anywhere in Michelangelo’s oeuvre. Out of 40 toes, all except for two fitted this brief: they had a short big toe and a long second toe, and the big toe goes outwards – it looks like someone is wearing a flip-flop in between the toes.”

He noted that the same toes can be seen on David and Moses in the Sistine Chapel in the Vatican, and added that other characteristics “shine through” in his work, including distinctive torsos and anatomically accurate pubic hair.

01:58 - Source: CNN
Moment Da Vinci painting sold for $450 million (2017)

“We all know a six-pack, but these guys actually have an eight-pack. I found two statues and five Michelangelo drawings that have that same, rare anomaly, which tells me that the model he used for those was the same model he used for these bronzes,” he said.

He also pointed to highly distinctive pubic hair that goes “up towards the umbilicus” in Michelangelo’s work, in contrast with the majority of classical and Renaissance sculptures, where pubic hair goes “down towards the genitalia.”

Abrahams’ conclusions were echoed by Victoria Avery, Keeper of Applied Arts at the Fitzwilliam Museum, who said the sculptures were the “real thing.”

“We believe these to be made by Michelangelo. And we think these are the originals made around 1505-1507,” she said, speaking at a press conference to announce the verification. “They are authentic Michelangelo, made when he was at the height of his creative genius, when he was desperate to outdo his contemporaries and dominate every medium on a massive scale.”

Professor Geraldine Johnson of the University of Oxford, an expert on Italian Renaissance sculpture, told CNN that the “groundbreaking” research carried out on the sculptures will force art historians to reconsider their assumptions of Michelangelo’s techniques.

Duane Tinkey/DSM Magazine
The "Apollo and Venus" painting by 16th-century Dutch master Otto van Veen (1556-1629) was discovered in the closet of an art gallery in Iowa, and is likely worth over $4 million.

Scroll through the gallery for other examples of lost and found artworks.
Bonhams
One of a triptych of artworks created by Ben Enwonwu during the aftermath of Nigeria's bloody civil war, "Tutu" disappeared shortly after being painted in 1974. Its whereabouts remained the subject of intense speculation for over 40 years before the portrait was discovered in a family home late last year. In March, it sold for over $1.6 million (£1,205,000).
Courtesy The Parker Gallery
"Two Hacks" (1789) by George Stubbs was sold at a Christie's "Living with Art" sale in New York in June 2016 -- originally listed as a copy. Art dealer Archie Parker -- believing it to be a real Stubbs -- purchased the painting for $175,000 ($215,000 with premium). The painting is currently hanging on his stand at the annual British Antique Dealers' Association (BADA) Fair in London, with an asking price of $900,000.
Courtesy: C.Cordes/Herzog Anton Ulrich Museum
Rembrandt's drawing of a dog has been in the collection of the Herzog Anton Ulrich Museum in Braunschweig, Germany, since 1770, but was long thought to be the work of a different artist.
Courtesy Tajan
In 2016, a drawing attributed to Italian master Leonardo da Vinci was discovered in Paris, after a portfolio of works was brought to Tajan auction house for valuation by a retired doctor. It was valued at 15 million euros ($16 million).
Courtesy Tajan
The drawing also features sketches of light and shadows and notes on the back.
Courtesy National Trust for Scotland
While researching for an episode of BBC's "Britain's Lost Masterpieces" series at the National Trust for Scotland's Haddo House collection in Aberdeenshire, art historian Bendor Grosvenor and a team of experts found a painting that could have been painted by artist Raphael.
Reuters
In April 2016, a painting believed to be by Caravaggio was found in an attic in France. Experts said it could be worth $136 million.
Courtesy National Trust for Scotland
The work was originally purchased for $25 dollars at the end of the 19th century. It could now be worth $26 million.
JEAN-PIERRE MULLER/AFP/Getty Images
In 1911, Leonardo Da Vinci's "Mona Lisa" was stolen from the Louvre by an Italian who had been a handyman for the museum. The famous painting was recovered two years later.
courtesy Art Recovery
A statue called "Young Girl with Serpent" by Auguste Rodin was stolen from a home in Beverly Hills, California, in 1991. It was returned after someone offered it on consignment to Christie's auction house. Rodin, a French sculptor considered by some aficionados to have been the father of modern sculpture, lived from 1840 until 1917. His most famous work, "The Thinker," shows a seated man with his chin on his hand.
AFP/Getty Images
Picasso's "La Coiffeuse" ("The Hairdresser") was discovered missing in 2001 and was recovered when it was shipped from Belgium to the United States in December 2014. The shipper said it was a $37 piece of art being sent to the United States as a Christmas present. The feds say it was actually a stolen Picasso, missing for more than a decade and worth millions of dollars.
ANDREAS SOLARO/AFP/Getty Images
Italy's Culture Ministry unveils two paintings by the French artists Paul Gauguin and Pierre Bonnard on April 2, 2014. The paintings, worth millions of euros, were stolen from a family house in London in 1970, abandoned on a train and then later sold at a lost-property auction, where a factory worker paid 45,000 Italian lire for them -- roughly equivalent to 22 euros ($30).
PAUL J. RICHARDS/AFP/Getty Images
A Renoir painting finished in the 1800s, loaned to a museum, reported stolen in 1951 and then bought at a flea market in 2010 has to be returned to the museum, a judge ruled on January 10, 2014. The 5½-by-9-inch painting, titled "Landscape on the Banks of the Seine," was bought for $7 at a flea market by a Virginia woman. The estimated value is between $75,000 and $100,000.
Courtesy Interpol
Seven famous paintings were stolen from the Kunsthal Museum in Rotterdam, Netherlands, in 2012, including Claude Monet's "Charing Cross Bridge, London." The paintings, in oil and watercolor, include Pablo Picasso's "Harlequin Head," Henri Matisse's "Reading Girl in White and Yellow," Lucian Freud's "Woman with Eyes Closed" and Claude Monet's "Waterloo Bridge," seen here. Works by Gauguin and Meyer de Haan were also taken.
New York County DA Office
Eight months after Salvador Dali's "Cartel de Don Juan Tenorio" was stolen in a New York gallery, a Greek national was indicted on a grand larceny charge in 2013.
Courtesy wga.hu
In 1473, Hans Memling's "The Last Judgment" was stolen by pirates and became the first documented art theft.
Hulton Archive/Getty Images
Adam Worth, the inspiration for Sir Arthur Conan Doyle's diabolical character Moriarty, stole "Georgiana, Duchess of Devonshire," painted by Thomas Gainsborough in 1876.
REUTERS/Chris Pizzello /Landov
Among their many crimes, the Nazis plundered precious artworks as they gained power during World War II. "Adele Bloch-Bauer I," by Austrian artist Gustav Klimt, was confiscated from the owner when he fled from Austria.
REUTERS /PHILIP SEARS /LANDOV
Many works of art that were taken by the Nazis were never recovered. Others were returned after years of legal battles. "Christ Carrying the Cross," by Italian artist Girolamo de' Romani, was returned to his family in 2012.
REUTERS/Handout /Landov
"The Scream" was one of two Edvard Munch paintings that were stolen from the Munch Museum in Oslo, Norway, in 2004.
REUTERS/Nelson Antoine /Landov
In 2007, Pablo Picasso's oil painting ''Portrait of Suzanne Bloch" was taken from the Sao Paulo Museum of Art. It was recovered two years later.

“Although we know from Renaissance sources that he also made sculptures in bronze, no surviving examples of works in this medium were believed to have survived,” she said.

The sculptures were first recorded as Michelangelo’s bronzes in 1878, when they were owned by the Rothschild family, but later lost their attribution.

The Fitzwilliam Museum subsequently claimed in 2015 that they were indeed Michelangelo originals.

The pair of sculptures was last sold at Sotheby’s in 2002, for around $2.3 million to a British collector, and was exhibited at the Royal Academy of Arts in London in 2012.